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    It is based on the principle that populations that share similar frequencies of a trait are more closely related than populations that have different frequencies of a trait. The genetic distance between two populations increases linearly with the geographic distance between them, due to isolation by distance and serial founder effects.

    When admixture occurs between populations, the genetic distance between the two populations is reduced. Cavalli-Sforza applied genetic distance measures to various populations around the world to infer phylogenetic relationships see Table on the right. All non-African populations are more closely related to one another i.

    This is consistent with a founder effect on the non-African population in that only a few individuals participated in the initial Out of Africa migration. The largest genetic distances observed are between Africa and Oceania and between Africa and the Americas. This is consistent with the isolation by distance and serial founder effects.

    Cavalli-Sforza suggests that the genetic distance between Sub-Saharan Africa and Europe is anomalously lower than it would be if the two populations had been evolving independently. The study suggests that the lower genetic distance between Europe and Africa can be explained by african dating site of europe admixture.

    Defining African admixture[ edit ] Generally, markers and lineages used to characterize African admixture are those that are believed to be specific to Africa. Ancient DNA analysis of these specimens indicates that they carried paternal haplotypes related to the E1b1b1b1a E-M81 subclade and the maternal haplogroups U6a and M1all of which are frequent among present-day communities in the Maghreb. These ancient individuals also bore an autochthonous Maghrebi genomic component that peaks among modern Berbersindicating that they were ancestral to populations in the area.

    Additionally, fossils excavated at the Kelif el Boroud site near Rabat were found to carry the broadly-distributed paternal haplogroup T-M as well as the maternal haplogroups K1T2 and X2the latter of which were common mtDNA lineages in Neolithic Europe and Anatolia. These ancient individuals likewise bore the Berber-associated Maghrebi genomic component.

    This altogether indicates african dating site of europe the Late Neolithic Kelif el Boroud inhabitants were ancestral to contemporary populations in the area, but also likely experienced gene flow from Europe. Y haplogroups R1and some paternal haplogroup T and U subclades. One subclade of haplogroup U, namely U6a1, is known to have expanded from northern and eastern Africa back into Europe [23] [24] even though haplogroup U6 is considered to have originated in the Middle East.

    Other lineages are known to have moved from Europe directly into Africa, for example mitochondrial haplogroups H1 and H3. Y-DNA[ edit ] One proposed example of Holocene gene flow from Africa to Europe, via the Middle East, is thought to be E1b1b 1, which is thought to have emerged about 22, years ago in East Africaand branches of it are thought to have migrated to the Middle East by 11, years ago during the late Pleistocene or early Neolithic periods.

    E1b1b1 lineages are found throughout Europe but are distributed along a south-to-north cline, with an E1b1b1a mode in the Balkans.

    In a sample of European males, Cruciani et al. The timing of this movement has been given widely varying estimates. During the Bantu Expansionpeople carrying haplogroup E not including E1b1b lineages dispersed across much of Sub-Saharan Africa from their homeland near the present-day border between Nigeria and Cameroon.

    The presence of E1b1a lineages outside Africa can typically be associated with events that occurred after the Bantu Expansion, such as the Moorish occupation of Iberia or the trade in African slaves. For example, Cruciani et al. E1b1a is closely related to E1b1b, the most frequent clade in Europe. E lineages that are not E1b1a or E1b1b could therefore reflect either a recent expansion associated with E1b1a or ancient population movements associated with E1b1b.

    The authors suggest that this distribution is consistent with a prehistoric migration from Africa to Iberia, possibly alongside mtDNA haplogroup U6. Haplogroups A and B are thought to have been the predominant haplogroups in central and southern Africa prior to the Bantu Expansion.

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    Currently these haplogroups are less common than E lineages. Haplogroup A has rare occurrences in Europe, but recently the haplogroup was detected in seven indigenous British males with the same Yorkshire surname. Another subclade, E3b1b-M81 is associated with Berber populations and is commonly found in regions that have had historical gene flow with northern Africa, such as the Iberian Peninsulathe Canary Islandsand Sicily. That is the highest frequency observed in Europe to date.

    According to a study in by Cerezo et al. In Iberia the mean frequency of haplogroup L lineages reaches 3. The authors suggest that this pattern indicates that most of the Sub-Saharan L african dating site of europe entered Iberia in prehistoric times rather than during the slave trade.

    According to Pereira et al.

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    They suggest this pattern is more compatible with a recent arrival of these lineages after slave trading began in the 15th century. The remains date to between the 12th and 13th centuries. The frequency of Sub-Saharan lineages detected in the medieval samples was The authors suggest the Muslim occupation and prehistoric migrations before the Muslim occupation would have been the source of these lineages.

    LatiumVolterra[44] Basilicataand Sicily. Though a high diversity of African mtDNA lineages have been detected, few lineages have accumulated enough mutations in Europe to form monophyletic clusters. L1b, L3b and L3d had matches with West African populations, indicating that these lineages probably entered Europe through Iberia. One lineage, L2a1a, appeared to be much older, indicating that it may have entered Europe in prehistoric times.

    This clade was possibly related to L2a1 clades identified in ten individuals of Ashkenazi heritage from France, Germany, Poland, Romania, and Russia. L2a lineages are widespread throughout Africa; as a result, the origins of this lineage are uncertain. The origins of haplogroup M1 have yet to be conclusively established. A study found that a prehistoric episode would be the main contributor to the sub-Saharan presence in Mediterranean Europe.
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